A study conducted by scientists in Brazil and the United Kingdom has quantified the impact that selective logging, partial destruction by burning, and fragmentation resulting from the development of pastures and plantations have had on the Amazon rainforest. In combination, these factors could be removing nearly 54 million tons of carbon from the forest each year, introduced into the atmosphere as greenhouse gases. This total represents up to 40 percent of the carbon loss caused by deforestation in the region.